Rapid Review·General Pathology

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INFLAMMATION

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Chemical Mediators of Inflammation (Arachidonic Acid Pathway)

P209

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The trunk: phospholipase A2 cleaves membrane phospholipids → arachidonic acid.
  • Inhibited by corticosteroids, which act at the very top and therefore shut off BOTH downstream arms.
  • This is the single most tested branch point on the page.
Arm 1, the COX pathway → prostaglandins and thromboxane. Inhibited by NSAIDs and aspirin.
  • Aspirin acetylates COX IRREVERSIBLY, so platelet inhibition lasts the ~7 to 10 day life of the platelet, which cannot make new enzyme. Other NSAIDs bind reversibly.
  • PGE2 and PGI2: vasodilation, ↑ vascular permeability, pain, fever.
  • PGE2 also protects the gastric mucosa (which is why NSAIDs cause ulcers, and why misoprostol, a PGE1 analog, prevents them) and maintains a patent ductus arteriosus (alprostadil keeps it open, indomethacin closes it).
  • Thromboxane A2, made by platelets: vasoconstriction plus platelet aggregation.
  • Prostacyclin (PGI2), made by endothelium: vasodilation plus inhibition of platelet aggregation. It is the exact counterweight to TXA2, and epoprostenol exploits it in pulmonary hypertension.
Arm 2, the LOX pathway (5-lipoxygenase) → leukotrienes. Inhibited by zileuton.
  • LTB4: potent neutrophil chemoattractant, alongside C5a and IL-8.
  • LTC4, LTD4, LTE4: bronchoconstriction, vasoconstriction, ↑ permeability. Together these are the "slow-reacting substance of anaphylaxis," blocked at the receptor by montelukast and zafirlukast.
  • Histamine from mast cells, basophils, and platelets: immediate vasodilation and permeability.
  • Bradykinin from kininogen via kallikrein: pain and permeability, and the cause of ACE-inhibitor cough and angioedema.
  • C3a and C5a, the anaphylatoxins; C5a is also chemotactic.
  • Platelet-activating factor (PAF) and NO (vasodilation via cGMP).

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What product of the arachidonic acid cascade promotes platelet aggregation and increases vascular tone?

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